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ƒA ƒJ ƒT ƒ^ ƒi ƒn ƒ} ƒ„ ƒ‰ ƒ –’Ϊ
ƒC ƒL ƒV ƒ` ƒj ƒq ƒ~ ƒŠ “‡ –’’θ
ƒE ƒN ƒX ƒc ƒk ƒt ƒ€ ƒ† ƒ‹ ’O –؏Ϊ
ƒG ƒP ƒZ ƒe ƒl ƒw ƒ ƒŒ
ƒI ƒR ƒ\ ƒg ƒm ƒz ƒ‚ ƒˆ ƒ

ƒG ( 233ƒŒƒR[ƒh)

˜a@–Ό‰Θ
ƒGƒCƒRƒEƒ„ƒbƒRƒ}ƒCƒ}ƒCƒAƒJƒ}ƒCƒ}ƒC’΄‰Θ ƒAƒJƒ}ƒCƒ}ƒC‰Θ
ƒGƒCƒRƒVƒ^ƒ_ƒ~ƒjƒVƒLƒEƒY’΄‰Θ ƒjƒVƒLƒEƒYƒKƒC‰Θ
ƒGƒKƒC ]ŠL (ƒjƒˆƒŠƒGƒKƒC)ƒtƒlƒKƒC’΄‰Θ ƒtƒlƒKƒC‰Θ
ƒGƒNƒAƒhƒ‹ƒgƒEƒKƒ^ƒ}ƒCƒ}ƒCƒgƒEƒKƒ^ƒ}ƒCƒ}ƒC’΄‰Θ ƒgƒEƒKƒ^ƒ}ƒCƒ}ƒC‰Θ
ƒGƒNƒAƒhƒ‹ƒ}ƒNƒ‰ƒAƒNƒLƒKƒC’΄‰Θ ƒ}ƒNƒ‰ƒKƒC‰Θ
ƒGƒNƒTƒ‰[ƒ^ƒMƒZƒ‹‚ΜˆŸŽνƒLƒZƒ‹ƒKƒC’΄‰Θ ƒLƒZƒ‹ƒKƒC‰Θ
ƒGƒNƒXƒpƒ“ƒTƒXƒkƒ}ƒKƒC (‰ΌΜ)ƒCƒVƒKƒC’΄‰Θ HYRIIDAE
ƒGƒWƒvƒgƒ_ƒJƒ‰ƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒ^ƒJƒ‰ƒKƒC‰Θ
ƒGƒXƒJƒŠƒtƒKƒ‰ƒpƒSƒXƒ}ƒCƒ}ƒC (‰ΌΜ)ƒgƒEƒKƒ^ƒ}ƒCƒ}ƒC’΄‰Θ ƒgƒEƒKƒ^ƒ}ƒCƒ}ƒC‰Θ
ƒGƒXƒJƒ‹ƒSƒ}ƒCƒ}ƒC’΄‰Θ ƒGƒXƒJƒ‹ƒS(ƒ}ƒCƒ}ƒC)‰Θ
ƒGƒXƒRƒoƒ‹ƒ{ƒ^ƒ‹ƒAƒNƒLƒKƒC’΄‰Θ ƒ}ƒNƒ‰ƒKƒC‰Θ
ƒGƒ]ƒCƒKƒCƒCƒKƒC’΄‰Θ ƒCƒKƒC‰Θ
ƒGƒ]ƒCƒOƒ`ƒCƒ‚ƒKƒC’΄‰Θ ƒNƒ_ƒ}ƒLƒKƒC‰Θ
ƒGƒ]ƒCƒVƒJƒQƒKƒCƒUƒ‹ƒKƒC’΄‰Θ ƒUƒ‹ƒKƒC‰Θ
ƒGƒ]ƒCƒ\ƒVƒWƒ~ƒjƒbƒRƒEƒKƒC’΄‰Θ ƒVƒIƒTƒUƒiƒ~ƒKƒC‰Θ
ƒGƒ]ƒCƒ\ƒjƒi ‰ΪˆΞˆι吃AƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒGƒ]ƒIƒIƒmƒKƒCƒIƒIƒmƒKƒC’΄‰Θ ƒIƒIƒmƒKƒC‰Θ
ƒGƒ]ƒKƒL (ƒiƒKƒKƒL)ƒJƒL’΄‰Θ ƒCƒ^ƒ{ƒKƒL‰Θ
ƒGƒ]ƒLƒŠƒKƒCƒ_ƒ}ƒVƒIƒjƒmƒcƒmƒKƒC’΄‰Θ ƒLƒŠƒKƒCƒ_ƒ}ƒV‰Θ
ƒGƒ]ƒLƒ“ƒ`ƒƒƒNƒCƒ^ƒ„ƒKƒC’΄‰Θ ƒCƒ^ƒ„ƒKƒC‰Θ
ƒGƒ]ƒRƒMƒZƒ‹ƒLƒZƒ‹ƒKƒC’΄‰Θ ƒLƒZƒ‹ƒKƒC‰Θ
ƒGƒ]ƒSƒ}ƒKƒC ‰ΪˆΞŒΣ–ƒŠLƒ„ƒ}ƒ^ƒjƒV’΄‰Θ ƒSƒ}ƒKƒC‰Θ
ƒGƒ]ƒSƒƒ‚ƒRƒƒ‚ƒKƒC’΄‰Θ ƒRƒƒ‚ƒKƒC‰Θ
ƒGƒ]ƒUƒ“ƒVƒ‡ƒE (ƒGƒ]ƒTƒ“ƒVƒ‡ƒE)ƒjƒVƒLƒEƒY’΄‰Θ ƒŠƒ…ƒEƒeƒ“ƒTƒUƒG‰Θ
ƒGƒ]ƒVƒ^ƒ_ƒ~ƒjƒVƒLƒEƒY’΄‰Θ ƒjƒVƒLƒEƒYƒKƒC‰Θ
ƒGƒ]ƒVƒ‰ƒIƒKƒCƒ‚ƒVƒIƒKƒC’΄‰Θ ƒGƒ]ƒVƒ‰ƒIƒKƒC‰Θ
ƒGƒ]ƒVƒƒoƒCƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒGƒ]ƒ^ƒ}ƒKƒCƒ^ƒ}ƒKƒC’΄‰Θ ƒ^ƒ}ƒKƒC‰Θ
ƒGƒ]ƒ^ƒ}ƒLƒIƒIƒVƒ‰ƒXƒiƒKƒC’΄‰Θ ƒ^ƒ}ƒLƒKƒC‰Θ
ƒGƒ]ƒ^ƒ}ƒLƒrƒ^ƒ}ƒLƒr’΄‰Θ ƒ^ƒ}ƒLƒrƒKƒC‰Θ
ƒGƒ]ƒ`ƒaƒ~ƒ{ƒ‰ƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒGƒ]ƒcƒmƒIƒŠƒCƒŒ ‰ΪˆΞŠpά“όƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒGƒ]ƒkƒmƒƒAƒTƒŠƒ}ƒ‹ƒXƒ_ƒŒƒKƒC’΄‰Θ ƒ}ƒ‹ƒXƒ_ƒŒƒKƒC‰Θ
ƒGƒ]ƒmƒnƒiƒKƒTƒcƒ^ƒmƒnƒKƒC’΄‰Θ ƒ†ƒLƒmƒJƒTƒKƒC‰Θ
ƒGƒ]ƒnƒ}ƒOƒŠ (ƒAƒjƒƒnƒ}ƒOƒŠ)ƒ}ƒ‹ƒXƒ_ƒŒƒKƒC’΄‰Θ ƒ}ƒ‹ƒXƒ_ƒŒƒKƒC‰Θ
ƒGƒ]ƒoƒC ‰ΪˆΞ?ƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒGƒ]ƒoƒJƒKƒCƒoƒJƒKƒC’΄‰Θ ƒoƒJƒKƒC‰Θ
ƒGƒ]ƒqƒ^ƒ`ƒIƒrƒAƒNƒLƒKƒC’΄‰Θ ƒKƒNƒtƒ{ƒ‰(ƒqƒ^ƒ`ƒIƒr)‰Θ
ƒGƒ]ƒqƒoƒŠƒCƒKƒC’΄‰Θ ƒCƒKƒC‰Θ
ƒGƒ]ƒqƒƒMƒZƒ‹ƒLƒZƒ‹ƒKƒC’΄‰Θ ƒLƒZƒ‹ƒKƒC‰Θ
ƒGƒ]ƒtƒlƒKƒCƒJƒŠƒoƒKƒTƒKƒC’΄‰Θ ƒJƒŠƒoƒKƒTƒKƒC‰Θ
ƒGƒ]ƒtƒlƒKƒC‚Μ’‡ŠΤƒJƒŠƒoƒKƒTƒKƒC’΄‰Θ ƒJƒŠƒoƒKƒTƒKƒC‰Θ
ƒGƒ]ƒ{ƒ‰ ‰ΪˆΞ–@—†ƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒGƒ]ƒ{ƒ‰ƒ‚ƒhƒL ‹[‰ΪˆΞ–@—†ƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒGƒ]ƒ}ƒCƒ}ƒCƒ}ƒCƒ}ƒC’΄‰Θ ƒIƒiƒWƒ}ƒCƒ}ƒC‰Θ
ƒGƒ]ƒ}ƒeƒ}ƒeƒKƒC‰Θ
ƒGƒ]ƒ}ƒƒ^ƒjƒV‚Μ’‡ŠΤƒŠƒ\ƒcƒ{’΄‰Θ ƒGƒ]ƒ}ƒƒ^ƒjƒV‰Θ
ƒGƒ]ƒ}ƒƒ^ƒjƒV‚Μ’‡ŠΤ02ƒŠƒ\ƒcƒ{’΄‰Θ ƒGƒ]ƒ}ƒƒ^ƒjƒV‰Θ
ƒGƒ]ƒ}ƒƒ^ƒjƒV‚Μ’‡ŠΤ03ƒŠƒ\ƒcƒ{’΄‰Θ ƒGƒ]ƒ}ƒƒ^ƒjƒV‰Θ
ƒGƒ]ƒ~ƒWƒ“ƒ}ƒCƒ}ƒCƒTƒiƒMƒKƒC’΄‰Θ ƒ~ƒWƒ“ƒ}ƒCƒ}ƒC‰Θ
ƒGƒ]ƒˆƒEƒ‰ƒNƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒGƒ]ƒƒXƒŒƒ}ƒ‹ƒXƒ_ƒŒƒKƒC’΄‰Θ ƒ}ƒ‹ƒXƒ_ƒŒƒKƒC‰Θ
ƒGƒ_ƒJƒ‰ƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒ^ƒJƒ‰ƒKƒC‰Θ
ƒGƒ`ƒIƒsƒAƒkƒ}ƒKƒC (‰ΌΜ)ƒCƒVƒKƒC’΄‰Θ HYRIIDAE
ƒGƒ`ƒ[ƒ“ƒrƒƒEƒhƒ}ƒCƒ}ƒCƒiƒ“ƒoƒ“ƒ}ƒCƒ}ƒC’΄‰Θ ƒiƒ“ƒoƒ“ƒ}ƒCƒ}ƒC‰Θ
ƒGƒbƒ`ƒ…ƒEƒoƒCƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒGƒfƒBƒLƒ…ƒ‰ƒˆƒEƒ‰ƒN (‰ΌΜ)ƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒGƒfƒ“ƒCƒ‚ƒCƒ‚ƒKƒC’΄‰Θ ƒCƒ‚ƒKƒC‰Θ
ƒGƒhƒCƒgƒJƒP ]ŒΛŽ…Š|ƒAƒTƒKƒIƒKƒC’΄‰Θ ƒCƒgƒJƒPƒKƒC‰Θ
ƒGƒhƒKƒƒ~ƒYƒSƒ}ƒcƒ{ (ƒEƒ~ƒSƒ}ƒcƒ{)ƒŠƒ\ƒcƒ{’΄‰Θ ƒ~ƒYƒSƒ}ƒcƒ{‰Θ
ƒGƒhƒ[ƒhƒxƒjƒAƒtƒŠƒJƒ}ƒCƒ}ƒC (‰ΌΜ)ƒAƒtƒŠƒJƒ}ƒCƒ}ƒC’΄‰Θ ƒAƒtƒŠƒJƒ}ƒCƒ}ƒC‰Θ
ƒGƒhƒ[ƒhƒˆƒEƒ‰ƒNƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒGƒiƒKƒVƒƒƒNƒVƒXƒiƒƒKƒC’΄‰Θ ƒVƒƒƒNƒVƒKƒC‰Θ
ƒGƒiƒKƒƒEƒoƒC (ƒ^ƒ“ƒZƒCƒGƒiƒKƒƒEƒoƒC)ƒVƒƒƒEƒoƒC’΄‰Θ ƒVƒƒƒEƒoƒC(ƒƒEƒoƒCƒKƒC)‰Θ
ƒGƒiƒƒ‹ƒAƒ}ƒKƒCƒAƒ}ƒIƒuƒl’΄‰Θ ƒAƒ}ƒIƒuƒlƒKƒC‰Θ
ƒGƒiƒƒ‹ƒXƒKƒCƒjƒVƒLƒEƒY’΄‰Θ ƒŠƒ…ƒEƒeƒ“ƒTƒUƒG‰Θ
ƒGƒkƒJƒ“ƒcƒuƒŠƒ{ƒ‰ (‰ΌΜ)ƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒGƒqƒƒ{ƒ^ƒ‹ƒAƒNƒLƒKƒC’΄‰Θ ƒ}ƒNƒ‰ƒKƒC‰Θ
ƒGƒrƒKƒC ŠC˜VŠLƒIƒjƒmƒcƒmƒKƒC’΄‰Θ ƒIƒjƒmƒcƒmƒKƒC‰Θ
ƒGƒrƒXƒKƒCƒjƒVƒLƒEƒY’΄‰Θ ƒjƒVƒLƒEƒYƒKƒC‰Θ
ƒGƒrƒXƒKƒC‚ΜˆκŽνƒjƒVƒLƒEƒY’΄‰Θ ƒjƒVƒLƒEƒYƒKƒC‰Θ
ƒGƒrƒXƒKƒC‚ΜˆκŽν ƒJƒŠƒuŠCƒjƒVƒLƒEƒY’΄‰Θ ƒjƒVƒLƒEƒYƒKƒC‰Θ
ƒGƒrƒ`ƒƒƒIƒŠƒCƒŒƒ{ƒ‰ƒRƒƒ‚ƒKƒC’΄‰Θ ƒRƒƒ‚ƒKƒC‰Θ
ƒGƒxƒŒƒbƒeƒBƒ‰ƒbƒpƒKƒC i‰ΌΜjƒTƒiƒMƒKƒC’΄‰Θ ƒLƒoƒTƒiƒMƒKƒC‰Θ
ƒGƒ}ƒCƒgƒLƒƒ„ƒcƒVƒƒKƒC’΄‰Θ ƒ„ƒcƒVƒƒKƒC‰Θ
ƒGƒ}ƒCƒ{ƒ^ƒ“ƒUƒ‹ƒKƒC’΄‰Θ ƒUƒ‹ƒKƒC‰Θ
ƒGƒ}[ƒ\ƒ“ƒˆƒEƒ‰ƒN (‰ΌΜ)ƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒGƒ~ƒpƒEƒ‹ƒXƒˆƒEƒ‰ƒN (‰ΌΜ)ƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒGƒ~ƒ}ƒNƒ‰ƒAƒNƒLƒKƒC’΄‰Θ ƒ}ƒNƒ‰ƒKƒC‰Θ
ƒGƒ~ƒ“ƒJƒƒzƒgƒgƒMƒXƒKƒC i‰ΌΜjƒGƒ[ƒŠƒAƒKƒC’΄‰Θ MUTELIDAE‰Θ
ƒGƒƒ‰ƒ‹ƒhƒ^ƒ}ƒSƒ}ƒCƒ}ƒCƒ}ƒCƒ}ƒC’΄‰Θ ƒIƒiƒWƒ}ƒCƒ}ƒC‰Θ
ƒGƒ‰ƒuƒ}ƒCƒ}ƒCƒ}ƒCƒ}ƒC’΄‰Θ ƒIƒiƒWƒ}ƒCƒ}ƒC‰Θ
ƒGƒŠƒUƒxƒXƒNƒ`ƒqƒŒ (‰ΌΜ)ƒ^ƒ}ƒLƒr’΄‰Θ ƒNƒ`ƒqƒŒƒKƒC‰Θ
ƒGƒŠƒUƒxƒXƒ{ƒ‰ƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒGƒŠƒUƒxƒXƒ{ƒ‰‚Μ’‡ŠΤƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒGƒŠƒUƒxƒXƒ{ƒ‰‚Μ’‡ŠΤ02ƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒGƒŠƒUƒxƒXƒ{ƒ‰‚Μ’‡ŠΤ03ƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒGƒŠƒbƒNƒXƒ`ƒhƒŠƒ}ƒXƒIƒoƒJƒKƒC’΄‰Θ ƒ`ƒhƒŠƒ}ƒXƒI‰Θ
ƒGƒŠƒ}ƒLƒ^ƒ}ƒ^ƒ}ƒKƒC’΄‰Θ ƒ^ƒ}ƒKƒC‰Θ
ƒGƒŠ[ƒ[ƒCƒ‚ƒCƒ‚ƒKƒC’΄‰Θ ƒCƒ‚ƒKƒC‰Θ
ƒGƒ‹ƒZƒ^ƒ€ƒ‰ƒTƒLƒnƒ‰ƒ_ƒJƒ‰ (‰ΌΜ)ƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒ^ƒJƒ‰ƒKƒC‰Θ
ƒGƒ‹ƒhƒ‰[ƒhƒGƒrƒXƒjƒVƒLƒEƒY’΄‰Θ ƒjƒVƒLƒEƒYƒKƒC‰Θ
ƒGƒ‹ƒxƒŠƒMƒZƒ‹ (ƒGƒ‹ƒxƒ‹ƒMƒZƒ‹)ƒLƒZƒ‹ƒKƒC’΄‰Θ ƒLƒZƒ‹ƒKƒC‰Θ
ƒGƒŒƒiƒ\ƒfƒVƒƒƒEƒoƒC’΄‰Θ ƒVƒƒƒEƒoƒC(ƒƒEƒoƒCƒKƒC)‰Θ
ƒGƒŒƒiƒcƒuƒŠƒ{ƒ‰ƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒGƒŒƒxƒbƒgƒEƒRƒ“ƒ}ƒCƒ}ƒCƒiƒ“ƒoƒ“ƒ}ƒCƒ}ƒC’΄‰Θ ƒiƒ“ƒoƒ“ƒ}ƒCƒ}ƒC‰Θ
ƒGƒŒƒxƒbƒgƒjƒWƒJƒPƒ}ƒCƒ}ƒCƒRƒEƒ‰ƒiƒƒNƒW’΄‰Θ ƒ}ƒ‰ƒbƒJƒxƒbƒRƒEƒ}ƒCƒ}ƒC‰Θ
ƒGƒŒƒ‰ƒIƒIƒ^ƒƒ‰ (‰ΌΜ)ƒ|ƒŠƒlƒVƒAƒ}ƒCƒ}ƒC’΄‰Θ ƒIƒIƒ^ƒƒ‰ƒKƒC‰Θ
ƒGƒƒ“ƒK[ƒ^ƒ~ƒNƒŠ (‰ΌΜ)ƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒGƒƒ“ƒ_ƒMƒZƒ‹ƒLƒZƒ‹ƒKƒC’΄‰Θ ƒLƒZƒ‹ƒKƒC‰Θ
ƒGƒ“ƒKƒmƒ}ƒŒƒCƒ}ƒCƒ}ƒCƒiƒ“ƒoƒ“ƒ}ƒCƒ}ƒC’΄‰Θ ƒiƒ“ƒoƒ“ƒ}ƒCƒ}ƒC‰Θ
ƒGƒ“ƒLƒ…ƒEƒVƒƒƒNƒV š’‹…ŽΫ”ΈƒXƒiƒƒKƒC’΄‰Θ ƒVƒƒƒNƒVƒKƒC‰Θ
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