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˜a–Ό‚Ε‚³‚ͺ‚·

ƒA ƒJ ƒT ƒ^ ƒi ƒn ƒ} ƒ„ ƒ‰ ƒ –’Ϊ
ƒC ƒL ƒV ƒ` ƒj ƒq ƒ~ ƒŠ “‡ –’’θ
ƒE ƒN ƒX ƒc ƒk ƒt ƒ€ ƒ† ƒ‹ ’O –؏Ϊ
ƒG ƒP ƒZ ƒe ƒl ƒw ƒ ƒŒ
ƒI ƒR ƒ\ ƒg ƒm ƒz ƒ‚ ƒˆ ƒ

ƒi ( 249ƒŒƒR[ƒh)

˜a@–Ό‰Θ
ƒiƒCƒYƒiƒLƒŠƒKƒCƒ_ƒ}ƒV (‰ΌΜ)ƒIƒjƒmƒcƒmƒKƒC’΄‰Θ ƒLƒŠƒKƒCƒ_ƒ}ƒV‰Θ
ƒiƒCƒ„ƒlƒ^ƒqƒ_ƒŠƒ}ƒLƒSƒ}ƒKƒC (‰ΌΜ)ƒ„ƒ}ƒ^ƒjƒV’΄‰Θ ƒSƒ}ƒKƒC‰Θ
ƒiƒJƒZƒRƒJƒƒjƒiƒIƒjƒmƒcƒmƒKƒC’΄‰Θ ƒJƒƒjƒi‰Θ
ƒiƒJƒ_ƒ`ƒMƒZƒ‹ƒLƒZƒ‹ƒKƒC’΄‰Θ ƒLƒZƒ‹ƒKƒC‰Θ
ƒiƒJƒ~ƒKƒƒJƒZƒ“ƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒiƒJƒ~ƒKƒƒJƒZƒ“‚ΜˆκŽνƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒiƒJƒ€ƒ‰ƒMƒZƒ‹ƒLƒZƒ‹ƒKƒC’΄‰Θ ƒLƒZƒ‹ƒKƒC‰Θ
ƒiƒJƒ„ƒXƒCƒ‚ ’†ˆΐˆπƒCƒ‚ƒKƒC’΄‰Θ ƒCƒ‚ƒKƒC‰Θ
ƒiƒJƒ„ƒXƒJƒZƒ“ƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒiƒJƒ„ƒ}ƒUƒ“ƒVƒ‡ƒEƒjƒVƒLƒEƒY’΄‰Θ ƒŠƒ…ƒEƒeƒ“ƒTƒUƒG‰Θ
ƒiƒKƒCƒVƒKƒC i‰ΌΜjƒCƒVƒKƒC’΄‰Θ MYCETOPODIDAE
ƒiƒKƒCƒgƒJƒPƒAƒTƒKƒIƒKƒC’΄‰Θ ƒCƒgƒJƒPƒKƒC‰Θ
ƒiƒKƒCƒgƒ}ƒLƒ{ƒ‰ƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒiƒKƒCƒ‚ (ƒTƒiƒ_ƒCƒ‚)ƒCƒ‚ƒKƒC’΄‰Θ ƒCƒ‚ƒKƒC‰Θ
ƒiƒKƒCƒ‚ƒtƒfƒAƒNƒLƒKƒC’΄‰Θ ƒtƒfƒKƒC‰Θ
ƒiƒKƒGƒ“ƒhƒEƒKƒCƒJƒŠƒoƒKƒTƒKƒC’΄‰Θ ƒqƒQƒ}ƒLƒiƒƒ{ƒ‰‰Θ
ƒiƒKƒIƒJƒ~ƒ~ƒKƒCƒIƒJƒ~ƒ~ƒKƒC’΄‰Θ ƒIƒJƒ~ƒ~ƒKƒC‰Θ
ƒiƒKƒIƒJƒ‚ƒmƒAƒ‰ƒKƒCƒIƒJƒ‚ƒmƒAƒ‰ƒKƒC’΄‰Θ ƒIƒJƒ‚ƒmƒAƒ‰ƒKƒC‰Θ
ƒiƒKƒIƒgƒƒKƒT (ƒAƒqƒ‹ƒmƒNƒ`ƒoƒV)ƒXƒJƒVƒKƒC’΄‰Θ ƒXƒJƒVƒKƒC‰Θ
ƒiƒKƒIƒŠ[ƒuƒcƒ„ƒ€ƒMƒ}ƒCƒ}ƒC (‰ΌΜ)ƒJƒTƒJƒ€ƒŠƒiƒƒNƒW’΄‰Θ ƒ„ƒ}ƒqƒ^ƒ`ƒIƒr‰Θ
ƒiƒKƒJƒYƒ‰ƒKƒCƒ„ƒcƒVƒƒKƒC’΄‰Θ ƒgƒEƒJƒ€ƒŠ‰Θ
ƒiƒKƒJƒmƒRƒ_ƒJƒ‰ƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒ^ƒJƒ‰ƒKƒC‰Θ
ƒiƒKƒJƒoƒtƒ_ƒJƒ‰ƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒ^ƒJƒ‰ƒKƒC‰Θ
ƒiƒKƒJƒƒ{ƒ^ƒ“ƒKƒCƒCƒVƒKƒC’΄‰Θ ƒCƒVƒKƒC‰Θ
ƒiƒKƒJƒƒ{ƒ^ƒ“ƒKƒC‚ΜˆŸŽνƒCƒVƒKƒC’΄‰Θ ƒCƒVƒKƒC‰Θ
ƒiƒKƒNƒrƒŒƒ}ƒCƒ}ƒCƒIƒIƒAƒJƒ}ƒCƒ}ƒC’΄‰Θ ƒIƒIƒAƒJƒ}ƒCƒ}ƒC‰Θ
ƒiƒKƒNƒƒIƒrƒ~ƒJƒ“ƒ}ƒCƒ}ƒC i‰ΌΜjƒgƒEƒKƒ^ƒ}ƒCƒ}ƒC’΄‰Θ ƒgƒEƒKƒ^ƒ}ƒCƒ}ƒC‰Θ
ƒiƒKƒNƒƒKƒlƒ_ƒJƒ‰ƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒ^ƒJƒ‰ƒKƒC‰Θ
ƒiƒKƒRƒoƒ“ƒXƒ\ƒLƒŒƒXƒJƒVƒKƒC’΄‰Θ ƒXƒJƒVƒKƒC‰Θ
ƒiƒKƒTƒLƒjƒVƒLƒjƒiƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒiƒKƒTƒ‰ƒTƒ~ƒiƒVƒCƒ‚ƒKƒC’΄‰Θ ƒCƒ‚ƒKƒC‰Θ
ƒiƒKƒTƒ‰ƒTƒ~ƒiƒV‚ΜˆκŒ^ƒCƒ‚ƒKƒC’΄‰Θ ƒCƒ‚ƒKƒC‰Θ
ƒiƒKƒUƒ‹ƒUƒ‹ƒKƒC’΄‰Θ ƒUƒ‹ƒKƒC‰Θ
ƒiƒKƒVƒ}ƒCƒ‚ƒCƒ‚ƒKƒC’΄‰Θ ƒCƒ‚ƒKƒC‰Θ
ƒiƒKƒVƒ}ƒMƒZƒ‹ƒLƒZƒ‹ƒKƒC’΄‰Θ ƒLƒZƒ‹ƒKƒC‰Θ
ƒiƒKƒVƒ}ƒ„ƒ^ƒeƒAƒNƒLƒKƒC’΄‰Θ ƒtƒfƒKƒC‰Θ
ƒiƒKƒVƒ‰ƒNƒTƒiƒKƒjƒVƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒiƒKƒXƒYƒJƒPƒ{ƒ‰ƒ„ƒcƒVƒƒKƒC’΄‰Θ ƒtƒWƒcƒKƒC‰Θ
ƒiƒKƒ\ƒEƒˆƒEƒCƒgƒJƒP –’ΪƒAƒTƒKƒIƒKƒC’΄‰Θ ƒCƒgƒJƒPƒKƒC‰Θ
ƒiƒKƒ^ƒPƒmƒRƒJƒjƒ‚ƒŠƒIƒjƒmƒcƒmƒKƒC’΄‰Θ ƒIƒjƒmƒcƒmƒKƒC‰Θ
ƒiƒKƒeƒ“ƒOƒKƒCƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒiƒKƒeƒ“ƒOƒjƒVƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒiƒKƒgƒMƒZƒ‹ƒLƒZƒ‹ƒKƒC’΄‰Θ ƒLƒZƒ‹ƒKƒC‰Θ
ƒiƒKƒgƒNƒTƒIƒJƒ`ƒ‡ƒEƒW i‰ΌΜjƒAƒtƒŠƒJƒ}ƒCƒ}ƒC’΄‰Θ ƒIƒJƒNƒ`ƒLƒŒ‰Θ
ƒiƒKƒiƒ^ƒlƒKƒCƒTƒiƒMƒKƒC’΄‰Θ ƒLƒoƒTƒiƒMƒKƒC‰Θ
ƒiƒKƒjƒVƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒiƒKƒjƒV—ή gŠCƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒiƒKƒkƒ}ƒ\ƒfƒKƒC cf.ƒVƒƒƒEƒoƒC’΄‰Θ ƒVƒƒƒEƒoƒC(ƒƒEƒoƒCƒKƒC)‰Θ
ƒiƒKƒmƒ‹ƒhƒW[ƒNƒAƒIƒMƒZƒ‹ i‰ΌΜjƒLƒZƒ‹ƒKƒC’΄‰Θ ƒLƒZƒ‹ƒKƒC‰Θ
ƒiƒKƒnƒcƒJƒlƒYƒ~ƒIƒjƒmƒcƒmƒKƒC’΄‰Θ ƒSƒ}ƒtƒjƒi‰Θ
ƒiƒKƒnƒiƒ}ƒ‹ƒ†ƒLƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒ^ƒJƒ‰ƒKƒC‰Θ
ƒiƒKƒoƒC ’·?ƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒiƒKƒqƒ_ƒgƒŠƒAƒIƒMƒZƒ‹ƒLƒZƒ‹ƒKƒC’΄‰Θ ƒLƒZƒ‹ƒKƒC‰Θ
ƒiƒKƒqƒƒlƒWƒKƒCƒAƒTƒKƒIƒKƒC’΄‰Θ ƒCƒgƒJƒPƒKƒC‰Θ
ƒiƒKƒqƒƒNƒ`ƒ_ƒJƒ‰ƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒ^ƒJƒ‰ƒKƒC‰Θ
ƒiƒKƒxƒbƒRƒEƒCƒ‚ƒCƒ‚ƒKƒC’΄‰Θ ƒCƒ‚ƒKƒC‰Θ
ƒiƒKƒzƒEƒIƒEƒCƒ‚ƒCƒ‚ƒKƒC’΄‰Θ ƒCƒ‚ƒKƒC‰Θ
ƒiƒKƒzƒŠƒJƒZƒ“ cf. ’·–x‰ΨεdƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒiƒKƒ~ƒmƒ€ƒVƒAƒNƒLƒKƒC’΄‰Θ ƒcƒNƒVƒKƒC‰Θ
ƒiƒKƒ~ƒ~ƒkƒ}ƒKƒCƒCƒVƒKƒC’΄‰Θ ƒCƒVƒKƒC‰Θ
ƒiƒKƒƒmƒEƒ_ƒJƒ‰ƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒ^ƒJƒ‰ƒKƒC‰Θ
ƒiƒKƒ†ƒEƒrƒVƒ‰ƒIƒ‚ƒVƒIƒKƒC’΄‰Θ ƒGƒ]ƒVƒ‰ƒIƒKƒC‰Θ
ƒiƒKƒŒƒKƒLƒJƒL’΄‰Θ ƒCƒ^ƒ{ƒKƒL‰Θ
ƒiƒKƒŒƒ{ƒŠƒVƒ‰ƒ^ƒ}ƒnƒiƒdƒgƒKƒC’΄‰Θ ƒVƒ‰ƒ^ƒ}ƒKƒC‰Θ
ƒiƒMƒTƒmƒVƒ^ƒ^ƒŠƒIƒJƒ~ƒ~ƒKƒC’΄‰Θ ƒIƒJƒ~ƒ~ƒKƒC‰Θ
ƒiƒMƒTƒ}ƒCƒ}ƒCƒŠƒ\ƒcƒ{’΄‰Θ ƒCƒ\ƒ}ƒCƒ}ƒC‰Θ
ƒiƒMƒiƒ^ƒ†ƒLƒ~ƒm ’·“α–ͺƒ~ƒmƒKƒC’΄‰Θ ƒ~ƒmƒKƒC‰Θ
ƒiƒTƒoƒC‚Μ’‡ŠΤ@“μƒAƒtƒŠƒJƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒiƒTƒoƒCƒcƒmƒ}ƒ^ƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒiƒVƒKƒ^ƒEƒ‰ƒVƒ}ƒ„ƒcƒVƒƒKƒC’΄‰Θ ƒgƒEƒJƒ€ƒŠ‰Θ
ƒiƒVƒKƒ^ƒ_ƒJƒ‰ (ƒCƒkƒrƒƒ_ƒJƒ‰)ƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒ^ƒJƒ‰ƒKƒC‰Θ
ƒiƒVƒKƒ^ƒ~ƒ„ƒVƒ —œŒ`‹{‘ツƒcƒVƒƒKƒC’΄‰Θ ƒ„ƒcƒVƒƒKƒC‰Θ
ƒiƒVƒWƒ_ƒJƒ‰ —œ’n•σƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒ^ƒJƒ‰ƒKƒC‰Θ
ƒiƒXƒCƒ‚ cf.ƒCƒ‚ƒKƒC’΄‰Θ ƒCƒ‚ƒKƒC‰Θ
ƒiƒXƒrƒoƒIƒgƒƒtƒf ‰ΦŽq—t‰³—•MƒAƒNƒLƒKƒC’΄‰Θ ƒcƒNƒVƒKƒC‰Θ
ƒiƒXƒ~ƒmƒ€ƒVƒAƒNƒLƒKƒC’΄‰Θ ƒcƒNƒVƒKƒC‰Θ
ƒiƒYƒ~ƒKƒC “D‚έŠLƒIƒJƒ~ƒ~ƒKƒC’΄‰Θ ƒIƒJƒ~ƒ~ƒKƒC‰Θ
ƒiƒ^ƒ}ƒƒMƒZƒ‹ ηλ“€‰ŒŠΗƒLƒZƒ‹ƒKƒC’΄‰Θ ƒLƒZƒ‹ƒKƒC‰Θ
ƒiƒ^[ƒ‹ƒAƒWƒƒCƒ‚ƒCƒ‚ƒKƒC’΄‰Θ ƒCƒ‚ƒKƒC‰Θ
ƒiƒ^[ƒ‹ƒAƒ‰ƒŒƒ^ƒ}ƒLƒrƒ^ƒ}ƒLƒr’΄‰Θ ƒ^ƒ}ƒLƒrƒKƒC‰Θ
ƒiƒ^[ƒ‹ƒCƒ‚ƒCƒ‚ƒKƒC’΄‰Θ ƒCƒ‚ƒKƒC‰Θ
ƒiƒ^[ƒ‹ƒJƒKƒ~ƒ}ƒ‹ƒXƒ_ƒŒƒKƒC’΄‰Θ ƒ}ƒ‹ƒXƒ_ƒŒƒKƒC‰Θ
ƒiƒ^[ƒ‹ƒJƒKƒ~‚Μ’‡ŠΤ?ƒ}ƒ‹ƒXƒ_ƒŒƒKƒC’΄‰Θ ƒ}ƒ‹ƒXƒ_ƒŒƒKƒC‰Θ
ƒiƒ^[ƒ‹ƒJƒ^ƒcƒmƒ}ƒ^ (‰ΌΜ)ƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒiƒ^[ƒ‹ƒRƒVƒ‰ƒ^ƒ} (‰ΌΜ)ƒnƒiƒdƒgƒKƒC’΄‰Θ ƒVƒ‰ƒ^ƒ}ƒKƒC‰Θ
ƒiƒ^[ƒ‹ƒTƒUƒGƒjƒVƒLƒEƒY’΄‰Θ ƒŠƒ…ƒEƒeƒ“ƒTƒUƒG‰Θ
ƒiƒ^[ƒ‹ƒVƒƒlƒYƒ~ƒVƒƒlƒYƒ~ƒKƒC’΄‰Θ ƒVƒƒlƒYƒ~ƒKƒC‰Θ
ƒiƒ^[ƒ‹ƒVƒƒnƒ}ƒOƒŠ (‰ΌΜ)ƒoƒJƒKƒC’΄‰Θ ƒoƒJƒKƒC‰Θ
ƒiƒ^[ƒ‹ƒXƒJƒVƒXƒJƒVƒKƒC’΄‰Θ ƒXƒJƒVƒKƒC‰Θ
ƒiƒ^[ƒ‹ƒzƒ^ƒ‹ƒoƒCƒAƒNƒLƒKƒC’΄‰Θ ƒ€ƒVƒƒKƒC‰Θ
ƒiƒ^[ƒ‹ƒˆƒEƒ‰ƒNƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒiƒbƒOƒXƒVƒ‚ƒuƒNƒŒƒ}ƒCƒ}ƒC (‰ΌΜ)ƒgƒEƒKƒ^ƒ}ƒCƒ}ƒC’΄‰Θ ƒgƒEƒKƒ^ƒ}ƒCƒ}ƒC‰Θ
ƒiƒbƒgƒCƒ‚ƒCƒ‚ƒKƒC’΄‰Θ ƒCƒ‚ƒKƒC‰Θ
ƒiƒcƒƒCƒ‚ ž₯ˆπƒCƒ‚ƒKƒC’΄‰Θ ƒCƒ‚ƒKƒC‰Θ
ƒiƒcƒƒGƒrƒX (ƒ^ƒ“ƒpƒGƒrƒX)ƒjƒVƒLƒEƒY’΄‰Θ ƒjƒVƒLƒEƒYƒKƒC‰Θ
ƒiƒcƒƒKƒC ž₯ŠLƒiƒcƒƒKƒC’΄‰Θ ƒiƒcƒƒKƒC‰Θ
ƒiƒcƒƒ_ƒJƒ‰ ž₯•σƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒ^ƒJƒ‰ƒKƒC‰Θ
ƒiƒcƒƒoƒC‚ΜˆκŽνƒAƒNƒLƒKƒC’΄‰Θ ƒ€ƒVƒƒKƒC‰Θ
ƒiƒcƒƒoƒC‚ΜˆκŽν02ƒAƒNƒLƒKƒC’΄‰Θ ƒ€ƒVƒƒKƒC‰Θ
ƒiƒcƒƒ{ƒ‰ƒAƒNƒLƒKƒC’΄‰Θ ƒKƒNƒtƒ{ƒ‰(ƒqƒ^ƒ`ƒIƒr)‰Θ
ƒiƒcƒƒ‚ƒhƒL ‹[ž₯•σƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒ^ƒJƒ‰ƒKƒC‰Θ
ƒiƒcƒ‚ƒ‚ —k”~ƒjƒVƒLƒEƒY’΄‰Θ ƒjƒVƒLƒEƒYƒKƒC‰Θ
ƒiƒcƒ‚ƒ‚—ήƒjƒVƒLƒEƒY’΄‰Θ ƒjƒVƒLƒEƒYƒKƒC‰Θ
ƒiƒfƒKƒ^ƒRƒIƒƒMƒ{ƒ‰ƒAƒNƒLƒKƒC’΄‰Θ ƒKƒNƒtƒ{ƒ‰(ƒqƒ^ƒ`ƒIƒr)‰Θ
ƒiƒfƒKƒ^ƒmƒVƒƒKƒ“ƒ[ƒLƒ{ƒ‰ƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒiƒfƒVƒRƒKƒC •ŽqŠLƒCƒ^ƒ„ƒKƒC’΄‰Θ ƒCƒ^ƒ„ƒKƒC‰Θ
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ƒiƒ“ƒˆƒEƒcƒLƒq “μ—mŒŽ“ϊŠLƒCƒ^ƒ„ƒKƒC’΄‰Θ ƒCƒ^ƒ„ƒKƒC‰Θ
ƒiƒ“ƒˆƒEƒiƒcƒƒ‚ƒhƒLƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒ^ƒJƒ‰ƒKƒC‰Θ
ƒiƒ“ƒˆƒEƒrƒi “μ—m—ƒAƒNƒLƒKƒC’΄‰Θ ƒ}ƒNƒ‰ƒKƒC‰Θ

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