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ƒA ƒJ ƒT ƒ^ ƒi ƒn ƒ} ƒ„ ƒ‰ ƒ –’Ϊ
ƒC ƒL ƒV ƒ` ƒj ƒq ƒ~ ƒŠ “‡ –’’θ
ƒE ƒN ƒX ƒc ƒk ƒt ƒ€ ƒ† ƒ‹ ’O –؏Ϊ
ƒG ƒP ƒZ ƒe ƒl ƒw ƒ ƒŒ
ƒI ƒR ƒ\ ƒg ƒm ƒz ƒ‚ ƒˆ ƒ

ƒc ( 256ƒŒƒR[ƒh)

˜a@–Ό‰Θ
ƒcƒ@[ƒ‰ƒAƒWƒƒCƒ‚ (‰ΌΜ)ƒCƒ‚ƒKƒC’΄‰Θ ƒCƒ‚ƒKƒC‰Θ
ƒcƒLƒKƒCƒcƒLƒKƒC’΄‰Θ ƒcƒLƒKƒC‰Θ
ƒcƒLƒKƒCƒ‚ƒhƒLƒcƒLƒKƒC’΄‰Θ ƒcƒLƒKƒC‰Θ
ƒcƒLƒqƒKƒC ŒŽ“ϊŠLƒCƒ^ƒ„ƒKƒC’΄‰Θ ƒCƒ^ƒ„ƒKƒC‰Θ
ƒcƒLƒˆƒmƒnƒ}ƒOƒŠƒ}ƒ‹ƒXƒ_ƒŒƒKƒC’΄‰Θ ƒ}ƒ‹ƒXƒ_ƒŒƒKƒC‰Θ
ƒcƒMƒmƒRƒ‚ƒ“ƒ_ƒJƒ‰ƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒ^ƒJƒ‰ƒKƒC‰Θ
ƒcƒMƒmƒXƒiƒMƒ“ƒ`ƒƒƒNƒCƒgƒJƒPƒAƒTƒKƒIƒKƒC’΄‰Θ ƒCƒgƒJƒPƒKƒC‰Θ
ƒcƒMƒmƒiƒKƒCƒgƒJƒPƒAƒTƒKƒIƒKƒC’΄‰Θ ƒCƒgƒJƒPƒKƒC‰Θ
ƒcƒNƒGƒKƒC ŠχŠLƒcƒNƒGƒKƒC‰Θ
ƒcƒNƒVƒKƒCƒAƒNƒLƒKƒC’΄‰Θ ƒcƒNƒVƒKƒC‰Θ
ƒcƒNƒVƒtƒ^ƒiƒVƒVƒƒƒWƒNƒCƒ‚ƒKƒC’΄‰Θ ƒNƒ_ƒ}ƒLƒKƒC‰Θ
ƒcƒNƒVƒ}ƒCƒ}ƒC ’}Ž‡ε—‹ƒ}ƒCƒ}ƒC’΄‰Θ ƒIƒiƒWƒ}ƒCƒ}ƒC‰Θ
ƒcƒOƒ`ƒKƒC ’ΓŒϋŠLƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒEƒ~ƒEƒTƒM‰Θ
ƒcƒQƒCƒ‚‚ΜˆκŽνƒCƒ‚ƒKƒC’΄‰Θ ƒCƒ‚ƒKƒC‰Θ
ƒcƒVƒ}ƒMƒZƒ‹ƒLƒZƒ‹ƒKƒC’΄‰Θ ƒLƒZƒ‹ƒKƒC‰Θ
ƒcƒVƒ}ƒPƒ}ƒCƒ}ƒCƒ}ƒCƒ}ƒC’΄‰Θ ƒIƒiƒWƒ}ƒCƒ}ƒC‰Θ
ƒcƒVƒ}ƒ}ƒCƒ}ƒCƒ}ƒCƒ}ƒC’΄‰Θ ƒIƒiƒWƒ}ƒCƒ}ƒC‰Θ
ƒcƒVƒ}ƒ„ƒ}ƒNƒ‹ƒ} ‘Ξ”nŽRŽΤƒ„ƒ}ƒ^ƒjƒV’΄‰Θ ƒ„ƒ}ƒ^ƒjƒV‰Θ
ƒcƒ^ƒmƒn ’Σ‚Μ—tƒcƒ^ƒmƒnƒKƒC’΄‰Θ ƒcƒ^ƒmƒnƒKƒC‰Θ
ƒcƒcƒJƒCƒRƒKƒCƒ_ƒ}ƒVƒLƒZƒƒ^ƒKƒC’΄‰Θ ƒXƒCƒtƒKƒC‰Θ
ƒcƒcƒJƒCƒRƒ_ƒ}ƒV‚Μ’‡ŠΤƒLƒZƒƒ^ƒKƒC’΄‰Θ ƒXƒCƒtƒKƒC‰Θ
ƒcƒcƒ\ƒfƒKƒCƒVƒƒƒEƒoƒC’΄‰Θ ƒVƒƒƒEƒoƒC(ƒƒEƒoƒCƒKƒC)‰Θ
ƒcƒcƒ~ƒKƒC •οŠLƒ^ƒ}ƒKƒC’΄‰Θ ƒ^ƒ}ƒKƒC‰Θ
ƒcƒdƒŒƒCƒgƒqƒLƒ}ƒCƒ}ƒCƒTƒ‰ƒTƒ}ƒCƒ}ƒC’΄‰Θ ƒTƒ‰ƒTƒ}ƒCƒ}ƒC‰Θ
ƒcƒdƒŒƒCƒ‚ƒCƒ‚ƒKƒC’΄‰Θ ƒCƒ‚ƒKƒC‰Θ
ƒcƒdƒŒƒLƒ“ƒ`ƒƒƒNƒCƒ^ƒ„ƒKƒC’΄‰Θ ƒCƒ^ƒ„ƒKƒC‰Θ
ƒcƒdƒŒƒgƒRƒuƒVƒIƒLƒiƒGƒrƒX’΄‰Θ ƒ~ƒ~ƒKƒC‰Θ
ƒcƒdƒŒƒiƒKƒLƒŠƒIƒŒƒ~ƒcƒNƒ`ƒLƒŠƒIƒŒƒKƒC’΄‰Θ ƒ~ƒcƒNƒ`ƒLƒŠƒIƒŒƒKƒC‰Θ
ƒcƒdƒŒƒiƒfƒVƒR ζAεψ•ŽqƒCƒ^ƒ„ƒKƒC’΄‰Θ ƒCƒ^ƒ„ƒKƒC‰Θ
ƒcƒdƒŒƒqƒƒJƒ^ƒxƒjƒVƒLƒEƒY’΄‰Θ ƒŠƒ…ƒEƒeƒ“ƒTƒUƒG‰Θ
ƒcƒdƒŒƒtƒfƒAƒNƒLƒKƒC’΄‰Θ ƒtƒfƒKƒC‰Θ
ƒcƒmƒCƒƒ`ƒ‡ƒEƒ`ƒ“ƒtƒfƒAƒNƒLƒKƒC’΄‰Θ ƒtƒfƒKƒC‰Θ
ƒcƒmƒCƒƒgƒŠƒmƒRƒAƒNƒLƒKƒC’΄‰Θ ƒwƒŠƒgƒŠƒKƒC‰Θ
ƒcƒmƒIƒŠƒCƒŒ cf. Špά“όƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒcƒmƒIƒŠƒCƒŒ‚Μ’‡ŠΤƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒcƒmƒIƒŠƒCƒŒ‚Μ’‡ŠΤ ƒAƒCƒXƒ‰ƒ“ƒhƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒcƒmƒKƒCƒcƒmƒKƒC‰Θ
ƒcƒmƒKƒC—ή –’ΪƒcƒmƒKƒC‰Θ
ƒcƒmƒKƒC—ή ‰«“κ –’ΪƒcƒmƒKƒC‰Θ
ƒcƒmƒLƒKƒC Šp–ΨŠLƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒcƒmƒLƒtƒfƒ}ƒNƒ‰ƒKƒCƒ‚ƒhƒL’΄‰Θ ƒGƒ]ƒcƒmƒ}ƒ^ƒKƒC‰Θ
ƒcƒmƒNƒ_ƒ}ƒLƒCƒ‚ƒKƒC’΄‰Θ ƒNƒ_ƒ}ƒLƒKƒC‰Θ
ƒcƒmƒNƒ_ƒ}ƒL‚Μ’‡ŠΤ01 –’ΪƒCƒ‚ƒKƒC’΄‰Θ ƒNƒ_ƒ}ƒLƒKƒC‰Θ
ƒcƒmƒRƒƒ‚ƒRƒƒ‚ƒKƒC’΄‰Θ ƒRƒƒ‚ƒKƒC‰Θ
ƒcƒmƒ_ƒVƒwƒiƒ^ƒŠ‚ΜˆκŒ^ƒIƒjƒmƒcƒmƒKƒC’΄‰Θ ƒEƒ~ƒjƒi‰Θ
ƒcƒmƒeƒcƒŒƒCƒV Šp“S?Ž}ƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒcƒmƒiƒKƒJƒZƒ“ƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒcƒmƒuƒGƒKƒCƒIƒjƒmƒcƒmƒKƒC’΄‰Θ ƒIƒjƒmƒcƒmƒKƒC‰Θ
ƒcƒmƒ}ƒ^ƒ‚ƒhƒL‚Μ’‡ŠΤƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒcƒmƒŒƒCƒVƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒcƒoƒCƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒcƒoƒTƒJƒmƒR cf.ƒAƒ}ƒIƒuƒl’΄‰Θ ƒAƒ}ƒIƒuƒlƒKƒC‰Θ
ƒcƒuƒJƒƒUƒ“ƒVƒ‡ƒEƒŠƒ\ƒcƒ{’΄‰Θ ƒJƒƒUƒ“ƒVƒ‡ƒEƒKƒC‰Θ
ƒcƒuƒRƒnƒNƒJƒmƒRƒAƒ}ƒIƒuƒl’΄‰Θ ƒVƒ‰ƒ^ƒ}ƒAƒ}ƒKƒC‰Θ
ƒcƒuƒŠƒ{ƒ‰ “ͺ–@—†ƒ„ƒcƒVƒƒKƒC’΄‰Θ ƒtƒWƒcƒKƒC‰Θ
ƒcƒuƒŠƒ{ƒ‰‚ΜˆκŽνƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒcƒuƒŠƒ{ƒ‰‚Μ’‡ŠΤƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒcƒ{ƒCƒ‚ƒCƒ‚ƒKƒC’΄‰Θ ƒCƒ‚ƒKƒC‰Θ
ƒcƒ{ƒEƒLƒdƒcƒEƒLƒdƒcƒKƒC‰Θ
ƒcƒ{ƒiƒŠƒRƒIƒƒMƒ{ƒ‰ƒAƒNƒLƒKƒC’΄‰Θ ƒKƒNƒtƒ{ƒ‰(ƒqƒ^ƒ`ƒIƒr)‰Θ
ƒcƒ{ƒiƒŠƒgƒŠƒmƒRƒAƒNƒLƒKƒC’΄‰Θ ƒwƒŠƒgƒŠƒKƒC‰Θ
ƒcƒ{ƒ~ƒLƒ‰ƒ‰ƒNƒ‹ƒ~ƒKƒC’΄‰Θ ƒNƒ‹ƒ~ƒKƒC‰Θ
ƒcƒ{ƒ~ƒŠƒ…ƒEƒOƒEƒUƒNƒ‰ƒjƒbƒRƒEƒKƒC’΄‰Θ ƒAƒTƒWƒKƒC‰Θ
ƒcƒ{ƒ~ƒƒ^ƒ]ƒRƒ€ƒVƒ –’ΪƒAƒNƒLƒKƒC’΄‰Θ ƒ€ƒVƒƒKƒC‰Θ
ƒcƒ}ƒOƒƒƒ“ƒlƒ‹ƒCƒ‚ƒCƒ‚ƒKƒC’΄‰Θ ƒCƒ‚ƒKƒC‰Θ
ƒcƒ}ƒOƒƒ„ƒiƒMƒVƒ{ƒŠƒ_ƒJƒ‰ƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒ^ƒJƒ‰ƒKƒC‰Θ
ƒcƒ}ƒWƒƒVƒYƒNƒjƒiƒCƒ‚ƒKƒC’΄‰Θ ƒNƒ_ƒ}ƒLƒKƒC‰Θ
ƒcƒ}ƒjƒPƒ{ƒŠ ελ’O–Ρ’€ƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒEƒ~ƒEƒTƒM‰Θ
ƒcƒ}ƒxƒjƒKƒCƒIƒIƒmƒKƒC’΄‰Θ ƒNƒ`ƒxƒjƒKƒC‰Θ
ƒcƒ}ƒxƒjƒNƒ_ƒ^ƒ}ƒKƒCƒLƒZƒƒ^ƒKƒC’΄‰Θ ƒXƒCƒtƒKƒC‰Θ
ƒcƒ}ƒxƒjƒqƒKƒC ƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒEƒ~ƒEƒTƒM‰Θ
ƒcƒ}ƒxƒjƒ}ƒcƒ€ƒV ελgΌ’ŽƒAƒNƒLƒKƒC’΄‰Θ ƒtƒgƒRƒƒKƒC‰Θ
ƒcƒ}ƒxƒjƒ}ƒƒAƒQƒ}ƒLƒEƒƒRƒKƒC’΄‰Θ ƒEƒƒRƒKƒC‰Θ
ƒcƒ}ƒxƒjƒƒ_ƒJƒ‰ƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒ^ƒJƒ‰ƒKƒC‰Θ
ƒcƒ}ƒ~ƒŠƒ…ƒEƒOƒEƒ{ƒ^ƒ‹ƒAƒNƒLƒKƒC’΄‰Θ ƒ}ƒNƒ‰ƒKƒC‰Θ
ƒcƒ}ƒ€ƒ‰ƒTƒLƒJƒZƒ“ƒAƒNƒLƒKƒC’΄‰Θ ƒAƒNƒLƒKƒC‰Θ
ƒcƒ}ƒ€ƒ‰ƒTƒLƒƒ_ƒJƒ‰ƒ^ƒJƒ‰ƒKƒC’΄‰Θ ƒ^ƒJƒ‰ƒKƒC‰Θ
ƒcƒ€ƒKƒ^ƒJƒjƒ‚ƒŠƒIƒjƒmƒcƒmƒKƒC’΄‰Θ ƒIƒjƒmƒcƒmƒKƒC‰Θ
ƒcƒ€ƒKƒ^ƒMƒZƒ‹ –aŽŒ^‰ŒŠΗƒLƒZƒ‹ƒKƒC’΄‰Θ ƒLƒZƒ‹ƒKƒC‰Θ
ƒcƒ€ƒKƒ^ƒRƒEƒVƒcƒuƒCƒ‚ƒKƒC’΄‰Θ ƒNƒ_ƒ}ƒLƒKƒC‰Θ
ƒcƒ€ƒKƒ^ƒRƒgƒcƒuƒCƒ‚ƒKƒC’΄‰Θ ƒNƒ_ƒ}ƒLƒKƒC‰Θ
ƒcƒ€ƒKƒ^ƒ\ƒfƒXƒCƒVƒ‡ƒEƒKƒC’΄‰Θ ƒXƒCƒVƒ‡ƒEƒKƒC(ƒ\ƒfƒ{ƒ‰)‰Θ
ƒcƒ€ƒKƒ^ƒ\ƒfƒ}ƒcƒ€ƒVƒAƒNƒLƒKƒC’΄‰Θ ƒtƒgƒRƒƒKƒC‰Θ
ƒcƒ€ƒKƒ^ƒ\ƒfƒ}ƒcƒ€ƒV‚Μ’‡ŠΤƒAƒNƒLƒKƒC’΄‰Θ ƒtƒgƒRƒƒKƒC‰Θ
ƒcƒ€ƒKƒ^ƒjƒiƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒcƒ€ƒKƒ^ƒlƒWƒ{ƒ‰ƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒcƒ€ƒKƒ^ƒmƒ~ƒMƒZƒ‹ƒLƒZƒ‹ƒKƒC’΄‰Θ ƒLƒZƒ‹ƒKƒC‰Θ
ƒcƒ€ƒKƒ^ƒƒƒ“ƒAƒNƒLƒKƒC’΄‰Θ ƒKƒNƒtƒ{ƒ‰(ƒqƒ^ƒ`ƒIƒr)‰Θ
ƒcƒ€ƒKƒ^ƒ‚ƒhƒLƒMƒZƒ‹ ‹[–aŽŒ^‰ŒŠΗƒLƒZƒ‹ƒKƒC’΄‰Θ ƒLƒZƒ‹ƒKƒC‰Θ
ƒcƒ€ƒoƒCƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒcƒ€ƒoƒC‚Μ’‡ŠΤ –’ΪƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒcƒƒ^ƒKƒCƒ^ƒ}ƒKƒC’΄‰Θ ƒ^ƒ}ƒKƒC‰Θ
ƒcƒƒiƒŠƒ~ƒ„ƒRƒhƒŠƒAƒ}ƒIƒuƒl’΄‰Θ ƒ†ƒLƒXƒYƒ‰Θ
ƒcƒ„ƒEƒ‰ƒVƒ}ƒ„ƒcƒVƒƒKƒC’΄‰Θ ƒgƒEƒJƒ€ƒŠ‰Θ
ƒcƒ„ƒGƒrƒXƒjƒVƒLƒEƒY’΄‰Θ ƒjƒVƒLƒEƒYƒKƒC‰Θ
ƒcƒ„ƒIƒIƒVƒ‰ƒ^ƒ}ƒnƒiƒdƒgƒKƒC’΄‰Θ ƒVƒ‰ƒ^ƒ}ƒKƒC‰Θ
ƒcƒ„ƒIƒiƒKƒ~ƒNƒŠƒAƒNƒLƒKƒC’΄‰Θ ƒGƒ]ƒoƒC‰Θ
ƒcƒ„ƒJƒTƒ}ƒCƒ}ƒCGASTRODONTOIDEA ƒJƒTƒ}ƒCƒ}ƒC‰Θ
ƒcƒ„ƒJƒXƒŠƒCƒ^ƒ„ƒCƒ^ƒ„ƒKƒC’΄‰Θ ƒCƒ^ƒ„ƒKƒC‰Θ
ƒcƒ„ƒKƒ‰ƒXƒCƒKƒC’΄‰Θ ƒCƒKƒC‰Θ
ƒcƒ„ƒLƒkƒ^ƒAƒQƒ}ƒLƒjƒbƒRƒEƒKƒC’΄‰Θ ƒLƒkƒ^ƒAƒQƒ}ƒL‰Θ
ƒcƒ„ƒMƒZƒ‹ƒLƒZƒ‹ƒKƒC’΄‰Θ ƒLƒZƒ‹ƒKƒC‰Θ
ƒcƒ„ƒNƒŒƒ^ƒAƒIƒMƒZƒ‹ (‰ΌΜ)ƒLƒZƒ‹ƒKƒC’΄‰Θ ƒLƒZƒ‹ƒKƒC‰Θ
ƒcƒ„ƒRƒKƒlƒ_ƒ}ƒAƒ}ƒIƒuƒl’΄‰Θ ƒ„ƒ}ƒLƒTƒS‰Θ
ƒcƒ„ƒRƒgƒcƒu ‰‹Υ¬—†ƒCƒ‚ƒKƒC’΄‰Θ ƒNƒ_ƒ}ƒLƒKƒC‰Θ
ƒcƒ„ƒRƒnƒNƒKƒC (‰ΌΜ)ƒRƒnƒNƒKƒC’΄‰Θ ƒRƒnƒNƒKƒC‰Θ
ƒcƒ„ƒTƒUƒGƒjƒVƒLƒEƒY’΄‰Θ ƒŠƒ…ƒEƒeƒ“ƒTƒUƒG‰Θ
ƒcƒ„ƒUƒNƒ‰ƒjƒbƒRƒEƒKƒC’΄‰Θ ƒjƒbƒRƒEƒKƒC‰Θ
ƒcƒ„ƒVƒCƒmƒ~ (ƒcƒ„ƒVƒCƒmƒ~ƒNƒ`ƒLƒŒ)ƒIƒIƒVƒCƒmƒ~ƒKƒC’΄‰Θ ƒIƒIƒVƒCƒmƒ~ƒKƒC‰Θ
ƒcƒ„ƒVƒƒƒWƒNƒCƒ‚ƒKƒC’΄‰Θ ƒNƒ_ƒ}ƒLƒKƒC‰Θ
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